The word snail is a common name for almost all members of the molluscan class Gastropoda that have coiled shells in the adult stage. When the word snail is used in a general sense, it includes sea snails, land snails and freshwater snails. Snails lacking a shell or having only a very small one are usually called slugs. Snails that have a broadly conical shell that is not coiled or appears not to be coiled are usually known as limpets.
Snails can be found in a wide range of environments from ditches, deserts, and the abyssal depths of the sea. Although most people are familiar with terrestrial snails, land snails are in the minority. Marine snails have much greater diversity and a greater biomass. The great majority of snail species are marine. Numerous kinds can be found in fresh water and even brackish water. Many snails are herbivorous, though a few land species and many marine species are omnivores or predatory carnivores.
Snails that respire using a lung belong to the group Pulmonata, while those with gills form a paraphyletic group; in other words, snails with gills are divided into a number of taxonomic groups that are not very closely related. Snails with lungs and with gills have diversified widely enough over geological time that a few species with gills can be found on land, numerous species with a lung can be found in freshwater, and a few species with a lung can be found in the sea.
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[edit] Land snails
Although the word snail is often used for all shelled gastropods, the word "snail" can also be used in a more limited sense to mean any of several species of large, air-breathing (pulmonate) land snails. Whichever land snail species is most commonly seen or most commonly eaten as escargot in a given area will usually be referred to as "snails" by the local people.
Species of land snails live in almost every kind of habitat, from deserts and mountains to marshes, woodland, and gardens. However, certain species are "anthropophilic", which means they are found most often around human habitation. Land snails have thinner shells, opposed to water snails, which sometimes have very thick shells.
[edit] Slugs
Gastropod species which lack a conspicuous shell are commonly called slugs rather than snails, although, other than having a reduced shell or no shell at all, there are really no appreciable differences between a slug and a snail except in habitat and behavior. A shell-less animal is much more maneuverable, and thus even quite large land slugs can take advantage of habitats or retreats with very little space – places that would be inaccessible to a similar-sized snail, such as under loose bark on trees or under stone slabs, logs or wooden boards lying on the ground.
Taxonomic families of land slugs and sea slugs occur within numerous larger taxonomic groups of shelled species. In other words, the reduction or loss of the shell has evolved many times independently within several very different lineages of gastropods, thus the various families of slugs are very often not closely related to one another.
[edit] Biology (primarily but not exclusively of pulmonate land snails)
[edit] Physical characteristics
Most snails move by gliding along on their muscular foot, which is lubricated with mucus and covered with epithelial cilia[1]. This motion is powered by succeeding waves of muscular contractions that move down the ventral of the foot. This muscular action is clearly visible when a snail is crawling on the glass of a window or aquarium. Snails move at a proverbially low speed (1 mm/s is a typical speed for adult Helix lucorum[2]). They produce mucus to aid locomotion by reducing friction, and the mucus also helps reduce the snail's risk of mechanical injury from sharp objects. This means that they can crawl along sharp objects like a straight razor and survive without injury.[3] Snails also have a mantle, a specialized layer of tissue which covers all of the internal organs as they are grouped together in the visceral mass, and the mantle also extends outward in flaps, which reach to the edge of the shell and in some cases can cover the shell, and which are partially retractable. The mantle is attached to the shell and creates it by secretion.
Most mollusks, including snails, have a shell which they have from the larval stage and which grows with them in size by the process of secreting calcium carbonate. Because of this, mollusks need calcium in their diet and environment to produce a strong shell. A lack of calcium, or low pH in their surroundings, can result in thin, cracked, or perforated shells. Usually a snail can repair damage to its shell over time if its living conditions improve, but severe damage can be fatal.
When retracted into their shells, many snails with gills (including many marine, some freshwater and some terrestrial species) are able to protect themselves with a door-like anatomical structure called an operculum. (The operculum of some sea snails has a pleasant scent when burned, so it is sometimes used as an ingredient in incense.)
Snails range greatly in size. The largest land snail is the Giant African Snail or Ghana Tiger Snail (Achatina achatina; Family Achatinidae), which can measure up to 30 cm. Pomacea maculata (Family Ampullariidae), or Giant Apple Snail is the largest freshwater snail, with a diameter of up to 15 cm and a mass of over 600 g. The biggest of all snails is Syrinx aruanus, an Australian marine species which can grow up to 77.2 cm (30 inches) in length[4] and 18 kg (40lbs) in weight.[5]
Most snails bear one or two pairs of tentacles on their heads. In most land snails the eyes are carried on the first (upper) set of tentacles (called ommatophores or more informally 'eye stalks') which are usually roughly 75% of the width of the eyes. The second (lower) set of tentacles act as olfactory organs. Both sets of tentacles are retractable in land snails. The eyes of most marine and freshwater snails are found at the base of the first set of tentacles.
[edit] Internal anatomy (land snails)
A snail breaks up its food using the radula, which is a chitinous structure containing microscopic hooks called cuticulae. With this the snail scrapes at food, which is then transferred to the digestive tract. This is why, in a quiet setting, a large land snail can be heard 'crunching' its food: the radula is tearing away at what it is eating.
The cerebral ganglia of the snail form a primitive brain divided into four sections. This structure is very much simpler than the brains of mammals, reptiles and birds, but nonetheless, snails are capable of associative learning.[6]
[edit] Growth of the shell
As the snail grows, so does its calcium carbonate shell. The shell grows additively, by the addition of new calcium carbonate, which is secreted by glands located in the snail's mantle. The new material is added to the edge of the shell aperture (the opening of the shell). Therefore the center of the shell's spiral was made when the snail was younger, and the outer part when the snail was older. When the snail reaches full adult size, it may build a thickened lip around the shell aperture. At this point the snail stops growing, and begins reproducing.
A snail's shell forms a logarithmic spiral. Most snail shells are right-handed, meaning that if the shell is held with the apex (the tip, or the juvenile whorls) pointing towards the observer, the spiral proceeds in a clockwise direction from the apex to the opening.
[edit] Hibernation/Estivation (land snails)
Some snails hibernate during the winter (typically October through April in the Northern Hemisphere). They may also estivate in the summer in drought conditions. To stay moist during hibernation, a snail seals its shell opening with a dry layer of mucus called an epiphragm.
Some freshwater snails such as apple snails have gills and a "door" or operculum to close the shell when they withdraw. This structure functions as protection from predators as well as protecting the soft tissues from desiccation when an aquatic habitat dries out temporarily.
[edit] Reproduction (primarily land snails)
All land snails are hermaphrodites, producing both spermatozoa and ova. Some freshwater snails, such as apple snails, and marine species, such as periwinkles, have separate sexes; they are male and female. The age of sexual maturity is variable depending on species of snail, ranging from as little as 6 weeks [7] to 5 years [8]. Adverse environmental conditions may delay the onset of sexual maturity in some snails [9].
Prior to reproduction, most land snails perform a ritual courtship before mating. This may last anywhere between two and twelve hours. Prolific breeders, pulmonate land snails inseminate each other in pairs to internally fertilize their ova. Each brood may consist of up to 100 eggs.
Pulmonate land snails and slugs have a reproductive opening on one side of the body, near the front, through which the outer reproductive organs are extruded so that exchange of sperm can take place. After this, fertilization occurs and the eggs develop.
Garden snails bury their eggs in shallow topsoil primarily while the weather is warm and damp, usually 5 to 10 cm down, digging with their foot. Egg sizes differ between species, from a 3 mm diameter in the grove snail to a 6 mm diameter in the Giant African Land Snail. After 2 to 4 weeks of favorable weather, these eggs hatch and the young emerge. Snails may lay eggs as often as once a month.
The snail's shell develops while it is still an embryo; it is, however, very weak, and needs an immediate supply of calcium. Newly hatched snails obtain this by eating the egg from which they hatched. The cannibalization by baby snails of other eggs, even unhatched ones, has been recorded. Promptly after they are finished ingesting their egg casings, they crawl upwards through the small tunnel in order to digest the egg. At this stage, the young are almost completely transparent and colorless. Their shell is usually slightly smaller than the egg they hatched from, but their length when out of their shell is slightly greater than the egg diameter. After a few weeks, the snails will begin to show their first tinge of color, usually slightly blue, before they turn their adult color. Roughly three months after they have hatched, they will look like miniature versions of their mature kin. They will continue to grow, usually for two to three years, until they reach adult size, although there have been confirmed recordings of snails growing amazingly fast - becoming even bigger than their parents in little more than a month.[citation needed] Irrespective of their rate of growth, however, it will still take at least 1 year before they are sexually mature.[10]
There have been hybridizations of snails; although these do not occur commonly in the wild, in captivity they can be coaxed into doing so.
Parthenogenesis has also been noted in certain species.[11][12]
Freshwater pond snails do not lay their eggs in the ground, but instead they attach them to something solid, or in some genera they carry the eggs internally until they hatch, a form of vivipary.
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